You could have mentioned changing the caps and checking the cables... I think the response from everyone would have been a little different.
The next step would be, for me, testing the optoisolators. I have attached a copy from one of the pages in the service manual for the 1281. There is a description of the serial path used to talk to the options boards.
I would also think about disconnecting all the boards, one at a time, and jumpering the links shown in the attached file. I haven't tried this myself but I think this would bypass each board and narrow down the cause of the error.