I'm not sure if it's common knowledge but it had me stumped for a bit to find a wire from a plate simply ending near the pre-amp section, connected to nothing.
That's what they are called because the wire "radiates" like 400Vpp at audio frequencies, as a gimmick capacitor with HF rolloff due to the 220k series resistor, in proximity to the sensitive pre-amp section.
Normally it does nothing but as soon as amplitude goes up and waveforms are a bit more square, there is some positive feedback and the amp's sound goes to a new dimension.
The video I'm not clear about what is the odd circuit he found, you don't need formal grid bias if you use "space-charge" instead. It's likely doing that, and when the adjacent triode upsets the space-charge then you get extra effects.
Musicians want a unique sound and get their amplifiers modded so no one else can sound like them.