Sure -- with an ohmmeter, probe across a pair of pads. It should read high or open (depending on which polarity you're using, there may be a small current through the logic chip, or the pullup resistors -- the straight carbon tracks). Pull the respective rubber piece from the top shell and try to position it over the board, and test the action. Whatever resistance it goes down to, is what it's doing.
You probably need about four hands to probe this, so take the time to do up connections carefully, or make a setup or something. Like, micro-clips could probe at the IC pins, hands-free; if you don't have any, you could solder jumpers onto respective pins, just for testing.
The resistance should be, Idunno, below a kohm or so? I forget what button pads usually do. In any case, it must be small in relation to the respective pullup resistance.
Board looks pristine, so I'd guess it's just button pad wear.
Tim