The replaced MC34151PG MOSFET driver in the original board suggests previous attempts to address potential driver problems. Both boards exhibit the LS1 error, indicating the fault might lie beyond the individual drivers.
You suspect a lack of signal to the MOSFET driver from the microcontroller, implying communication issues. Tracing the incline potentiometer to the microcontroller's POT output pin (12) is a good investigative step.
One more troubleshooting step: The incline sensor which is responsible for
speed & incline of Sole, is maybe working fine. But I want you to trace this sensor wire which is connecting in controller board. Remove the wire from board, take off the little white piece which helps inserting pin easily, and then insert without this headpiece. You can take help from the picture below, related to piece which you need to remove.
Picture 1: Incline sensor head(in skin colour) , separate this from actual pin by pulling out.
Picture 2: Demonstrating how it separates and showing 2 separate pieces.
Picture 3: Re-insert without the head.