They will be 5V 50mA lamps, so you can replace them with a LED and a series resistor in those places where it is fed with a constant voltage, and with a shunt resistor added in those places where they are in series and driven with a current. Tip for the LED is to use a red or orange LED for those with red or orange caps, and green for the green caps, with white for clear ( or choose a LED colour) and run them at under 5mA, otherwise they will be too bright.
I used to replace a lot of soldered in dial illumination lamps, so still have a small box of the assorted grain of wheat lamps around, in 5V, 12V and 28V varieties. As it is a mixed box I test before using, using a variable supply and a 10R resistor.
If you really cannot get one you can transplant a LED and a 1/8W resistor into those little lamp units after taking the glass out. difficult, but doable using a very small 3mm LED.