What you're seeing is known as Cathode poisoning. It is caused by contamination of the emmissive surface of the digit cathodes normally from sputtering of material from other, more used, digits. It is almost guaranteed for digits that are almost always blank.
Often poisoning is an indication of end-of-life. Even if the tube is an old NOS one, they still have a gas leakage rate past the pin seals. Atmospheric Helium is a particular problem (for vacuum tubes too) because it cannot be captured by the getters. I'm not sure if Helium leak rate is responsible for increased operating voltage.
Experimental restoration has varied success and methods. The gentlest is to keep all cathodes cycling at normal current, it takes a long time and is the recommended method of retaining normal operation. running at elevated current can help, Burroughs recommend a higher supply rather than reducing the anode resistor, this promotes easier striking.
The brave method is to run each digit in turn using a
significantly higher current. This sputters off material from the surface and reveals clean material. This needs to be done with caution, running each digit until it is fully illuminated and then move onto the next one. There is a danger of the sputtered material settling onto the other digit cathodes and poisoning them so sequencing them is important. The other danger is that, if you go too far, material can be sputtered onto the glass and the supporting insulators between the cathode element, causing blackening or leakage. [EDIT: Of course, you would return the Nixie to normal circuit operation after treatment. Borroughs recommended using the lowest acceptable brightness (current) for longest life - by high HT voltage, 175V or so, and higher value anode resistor].
I don't know if you have found the Borroughs Nixie databook, you can find it here if not...
https://frank.pocnet.net/other/Burroughs/616.pdfP.S. Some Soviet nixies, eg. the IN-1, omitted the trace mercury content, this lead to very short life with cathodes literally disintegrating and shorting. This won't be an issue with Borroughs displays though.